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What about the Bark Beetle?
(*Notes:
This article was first presented at the regular
meeting of the Botanical Society on January 6,
1996. A related article, written by Ed Berg,
appeared in the Peninsula Clarion in October 13,
2000 and can be viewed on
this page in their on-line
archives.
Also see additional plant nutrition
information in the nitrogen
article,
in the observations listings.
This
article was written originally about the spruce
forests of Southcentral Alaska, but the information
is applicable to ALL temperate evergreen
forests.)
How
Forests Grow
Below
is and illustration showing the stages of forest
development. You can simply scroll down and read
about each step in forest growth, or you can click
on the letters under the illustration (A - J) to
link to corresponding description in the article.
To return here, click on any of the red letters in
the article (A -J) .
A
-
In Alaska, the beginning of forest growth is when
glaciers recede leaving sterile scarred lands
consisting of sand, gravel and boulders. Few plant
species can live and colonize such nutritionally
poor environments. A few small hardy species can
colonize and they try; some seeds blown into
crevices and covered with wind-blown dust and fine
sand is enough for a start. Their life expectancy
is very short. As the plant grows, its demands for
more nutrition are not met, and these first
pioneers usually die, decompose, and create new and
richer soil for the next individual to grow in.
This continues throughout the glaciated area, some
places are more conducive to plant growth, with
more water holding qualities and perhaps on the
south side of a raised area. Other places might
remain barren for a thousand years. The first
plants are sure to die of starvation, leaving
behind composting material for the next generation
to live in. Each time a plant community thrives,
runs out of nutrients and dies, it adds to the
buildup of soil consisting of rich, composted
materials for a larger variety of diverse plant
species. But these too will ultimately succumb to
lack of nutrition and as they weaken, insects will
invade the colony hastening its demise.
The
problem facing plant growth is that sand and gravel
does not store water, they also allow the
nutrients, in what little top soil there is, to
leach out where it is lost. This situation worsens
in northern environments because the temperature a
short distance below the surface is too cold. Even
though an area might receive forty inches of
rainfall a year, if the water has nothing to keep
it out of the gravel, or unless the water table is
near the surface, drought can occur within a few
days of the last rain storm. Deep snow of winter is
not very helpful for watering plants either, this
too melts and seeps away. Only deep soils can stop
the loss of water, whether they are too cold for
root growth is not the issue, what is at issue is
water retention, something that will hold nutrients
for the plants to use between rains.
How
much soil does a plant require?
It is a
well known fact that nearly all terrestrial plants
(some epiphytic species glean most nutrients from
the air) must have equal amounts of shoot and root.
When one looks at a large tree it is amazing to
believe that an equal amount of that tree's mass is
below ground. That is why we must "thin out" garden
seedlings, giving each plant enough space for the
mass of roots which allows it to thrive. Whenever
soil is shallow, drought can have an immediate
effect. Crowding only hastens the problem, although
with some species, such as many kinds of grasses,
root mass can hold quite a bit of water but, roots
do not thrive without nutrition too The roots which
remove nutrients and water from the soil are not
the ones which are easily seen, they are minute
hair-like structures growing from small rootlets.
They are extremely vulnerable to drought in shallow
soil.
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Pictured
at right is a small plant with the amount
of root necessary to allow it to live, but
not enough to allow much more growth or
reproduction. If there are a number of
plants near each other, each one will
attempt to put out roots equal to the
shoot, and the end result is they quickly
use up all of the nutrients, and those
that live will not be able to sustain a
normal growth rate unless there is deep
soils and plenty of water to help compost
the dead plants. If the soil is shallow
the trees will never reach their optimal
height and width.
Now,
let us take the knowledge we have and move
on. The illustration at the top of this
page is not meant to be a panorama view,
rather it is a series of steps, from A to
J that a forest must go through on a
single piece of land, perhaps only a few
acres or miles in diameter. Many things
can effect it due to the presence of
hills, rivers, bog lakes, etc.
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B
-
As the soil is built up from dead plants, compost
is added, from the dead roots as well as the dead
upper plant parts. Willows and populus are often
the first larger woody plants to grow, and in some
areas, alders and many herbaceous species too
numerous to name here. As with the preceding growth
in A, these kill most of them, a few days of
drought will weaken them and they will succumb to
insect attack and surely die out. At this stage in
the forest's development the first spruce appear,
they can withstand more shade than deciduous
species and will live much longer, although stunted
and gnarled. Black spruce will persist for over one
hundred years in areas where white or Sitka spruce
cannot. But even these will add to the soil depth
as they are reduced by bacteria, fungi, insects,
and other natural composters.
In the
place where B was, now appears the first "mixed
woods".
C
- Within this stand there will be a greater number
of species of deciduous plants, mosses and broad
leafed evergreens, such as lingonberry, club mosses
horsetails, cornus and many more. These add compost
as they die as do the leaves of aspen, cottonwood,
birch and alder. The alders are very beneficial
because they have nitrogen fixing bacteria, and add
greatly to nitrogen poor soils. The spruce, being
protected by the faster growing deciduous species,
thrives and within time their numbers increase and
they grow into a dense shaded umbrella which kills
any species that must have abundant sunlight. In
fact the lower limbs of spruce, growing in dense
groves, will lose all of their lower limbs from
lack of sunlight.
D
- What has happened is, through natural forest
succession, a very small and weak climax spruce
forest has been born. This has been a long time
building, perhaps as much as a five hundred years.
But alas, the reason it is filled with odd shaped
spruce trees is simply that there is not enough
soil depth yet, and they have used up all of the
nutrients provided by the other dead plants. They
weaken and once again they are vulnerable to attack
by insects, usually bark beetles, psyllids, and a
host of others, including fungi in the form of
molds and rusts. This creates the next die
off.
E
-
The die off pictured is from bark beetles, a fire
is not pictured because it robs the forest of the
natural compost and nitrogen which is needed to
grow a better forest than the one just devastated.
Fires leave a great deal of carbon and other
chemicals needed for plant growth but, to grow in
what? The same shallow soil, and the forest is
literally "set back" about one hundred and fifty
years. If however, spruce trees do not grow too
soon and in great numbers, the forest can through
natural succession, once again build good composted
soil, and the next time the forest becomes climaxed
with spruce, they will be larger and
stronger.
F
- Is about where most of the Alaskan forests,
currently being devastated by the bark beetles are
at this time. Some are older and have gone through
an additional stage, and perhaps have had more
moisture. They are still a long way from being
capable of sustaining themselves against much
drought or any of a great number of conditions and
enemies.
G
- Natural succession occurs, and as the dead tree
roots decay and the trees fall to the ground, new
composting takes place and soil depth is again
added to the forest floor. The next forest is very
strong but still vulnerable but to a lesser
degree.
H
-
I
-
These too will be struck by beetles or other
natural composters, but their die off numbers will
be less and the recovery more swift.
J
- Represents a very old but strong forest, healthy,
but always with some bark beetles present. If
general weakness occurs within the forest the
beetles will return in force, causing another major
die off. The beetles will be there when the last
spruce tree fades into memory.
The
bark beetle is without a doubt the most important
insect in the growth of strong forests. As with
wolves and other predators, they prey upon the weak
and guarantee stronger generations to come. But all
of their work will be for nothing, if the dead
material is hauled away and not allowed to
decompose. All it will do is guarantee another weak
forest that beetles will be waiting for.
Note:
Dead fallen logs are water containers that assist
forests during dry periods, and furnish habitat for
voles which are important in the food chain, and
for moving mycorrhizal fungus spores, which are a
vital part of any forest.
--End
*See
Related story in the Peninsula
Clarion
Archives
by Ed Berg
Also see additional plant nutrition
information in the nitrogen
article,
in the Observation's articles.
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